CEDAR FALLS, Iowa (AP) — Northern Iowa kept three-time defending FCS champion North Dakota State out of the end zone and handed the Bison their first loss, 23-3 on Saturday.

The loss snapped North Dakota State’s 33-game winning streak, the longest in Football Championship Subdivision history. It also snapped an 18-game Missouri Valley Football Conference win streak. The loss knocked the Bison (9-1, 5-1) into a first-place tie with Illinois State atop the MVFC standings. Northern Iowa (6-4, 4-2) pulled into a tie with Youngstown State for second place.

David Johnson rushed for 133 yards on 28 carries to lead Northern Iowa. Brion Carnes finished with 43 yards, including a 6-yard run for a first-quarter touchdown. David Miller scored on a 30-yard run in the fourth quarter to seal the victory.

North Dakota State finished with just 175 yards of total offense and was just 1-for-14 in third down efficiency.